Genetic Topics

The Connemara Pony in South Africa

THE HISTORY OF THE CONNEMARA PONY IN SOUTH AFRICA

ZIMBABWE

Keirin of Leam by Sean of Leam out of Teresa of Leam, arrived in Zimbabwe in September 1968 imported by Miss Worthington in Zimbabwe. Keirin’s son Dundrum (CPS 800) born 1976, out of Coromandel Abbeyline (CPS 8118) bred by Sydney Press was subsequently acquired by Christine Walwyn. Both were inspected by Donal Kenny in 1985 and registrant with the Irish Connemara Society.

LESOTHO

In 1978 Milford Hurricane (CPS 568) (Mervyn Bowsprit – Mervyn Silver Grey) and Crockaun Connemara (also known as Croc on Oir, by the Fugitive – Silver Lace) were purchased for the Lesotho National Stud as the result of a project undertaken by Donal Kenny on behalf of the Irish Government in consultation with the Lesotho Government. The aim of the project was to conserve and improve the Basotho Pony upon whom the Basotho farmers relied heavily as a means for transport. As a result of the Boer War thousands of their best bred stock was lost and the Basotho Pony breed was sadly depleted. The Irish Government donated two stallions, Milford Hurricane and Croc an Oir. They were taken to the Lesotho National Stud at Thaba Tseka.

connemara_south_africa_2012_img_0

(L-R: Pictures Christine Walwyn, Progeny of Dundrum; Ashgar Ripley (PB winner of the Connemara Victor Ludorum HOY 2011)

At the same time, in Clarens (South Africa) on the Lesotho Border, Christine Walwyn’s first Connemara Pony Abbeyleix Tansey (CPS 3103) Carna Dun-Pretty Molly) dam of the Cornmandel Abbeyline was purchased by her from Sydney Press. Abbeyline and her daughter Keirina (CPS 119) were given to Mrs Walwyn by Mrs Worthington when Zimbabwe (Rhodesia) was experiencing difficult times. They travelled to safety to South Africa with the stallion Dundrum (CPS 800) these four ponies became the foundation of the Ashgar Stud whose aim was to provide children’s ponies for all disciplines. Later Mrs Walwyn leased Croc an Oir from the Basotho Pony Stud. After a perilous journey through the Lesotho Mountains he arrived safely at the Ashgar Stud where he sired Ashgar Doo Gary, Ashgar Donegal and Ashgar Karralee. These three ponies brought new bloodlines to the Ashgar Stud.

At last all three of the original bloodlines imported from different sources at different times and so widely came together at Ashgar Stud in the Free State.

Today, Ashgar Doo Gary and Ashgar Donegal are near retirement age and are resident at Ashgar Stud in Clarens. Ashgar has been represented in the junior showing and eventing teams throughout South Africa.

KILSHANE IRISH SPORT HORSE STUD, CAPE TOWN

L-R: Kilshane Christoper Robin (CPBS 97/511 Sire – Templebready Fear Bui, Dam Wild Winnie) ridden by Bronwyn Hughes, Johannesburg, Gauteng South Africa

L-R: Kilshane Christoper Robin (CPBS 97/511 Sire – Templebready Fear Bui, Dam Wild Winnie) ridden by Bronwyn Hughes, Johannesburg, Gauteng South Africa

In 1998 Ian and Bea Hurst moved to the Western Cape South Africa. They brought with them two stallions, Kilshane Clover (Irish Draught) and Kilshane Christopher Robin (1997 Connemara). The stud is now firmly established on their farm in the Franschoek district, where the broodmares and weanlings are kept. They currently breed with their two stallions and do not make use of imported semen. Kilshane Christopher Robin stands in Gauteng where he competes regularly in showing and eventing. He has won the Connemara Working Riding championship in 2010 as well as the In-hand Championship for Connemaras at the Horse of the Year Show in 2009.

Christopher Robin’s dam won several show classes, whilst his sire was winner of the 1993 and 1994 performance test programme run by the Irish Government. Christopher Robin’s pedigree shows the presence of bay lines, important in keeping hard colours and avoiding blue-eyed creams. He also shows the typical characteristics of his breed, having a super temperament, intelligence, hardiness, and good jumping ability. He has part-bred foals competing in the Cape, Free State and Gauteng areas of South Africa.

MILLSLEIGH IRISH SPORT HORSE AND PONY STUD, LADYBRAND, FREE STATE

L -R: Ashgar Ripley (Part-bred WB x Conn by Christopher Robin); Victor Ludorum Horse of the Year 2011; Millsleigh James Morrison (Part-bred Thb x Conn by Christopher Robin); Ashgar Ripley; Millsleigh Jonathan Jo (Part-bred Thb x Conn by Christopher Robin) Stallion: Ashgar Cruising CSSA 1004 (by Ashgar Donegal -Sire Croc on Nor, Dam Coromandel Abbeyline out of Dam Dun o’Clair – Sire Dundrum, Dam Abbeyleix Tansey)

L -R: Ashgar Ripley (Part-bred WB x Conn by Christopher Robin); Victor Ludorum Horse of the Year 2011; Millsleigh James Morrison (Part-bred Thb x Conn by Christopher Robin); Ashgar Ripley; Millsleigh Jonathan Jo (Part-bred Thb x Conn by Christopher Robin) Stallion: Ashgar Cruising CSSA 1004 (by Ashgar Donegal -Sire Croc on Nor, Dam Coromandel Abbeyline out of Dam Dun o’Clair – Sire Dundrum, Dam Abbeyleix Tansey)

Since 2005 Millsleigh Irish Sport Horse and Pony Stud has been a breeder and producer of quality young Sport Horses of Irish Draught lines (Stallions: Kinvara’s Etherow Impasse and Gleneagle Rebel) with proven Thoroughbred sport horses from Eventing, Show Jumping, Dressage and Polo disciplines. The foundation mares were and continue to be selected for their temperament, athleticism and proven record. Progeny are now aged between 9 months (including a surviving twin filly) and 5 years. After critical review of the mares’ strengths and weaknesses, the Kinvara, Ashgar and Kilshane stallions were chosen to compliment the mares’ temperaments, conformation and to add height, substance and quality. All progeny were bred by Artificial Insemination to Irish Draught and Connemara stallions based in other parts of the country which presents many logistical challenges. The strategy of the stud is to breed and produce Eventers, Working Hunters and Working Hunter Pony stars of the future with added social, easy going temperaments.

Ashgar Cruising a young stallion now five-years-old is the resident foundation stallion at the Millsleigh Irish Sport and Pony Stud (owned by Dipereng Equestrian Services) near Ladybrand in the Free State Province. 2011-12 is his first breeding season, with two pure-bred Ashgar mares by Dundrum, Ashgar Felipe and Gaymaree and several thoroughbred mares. We are looking forward to seeing his first pure and part-bred progeny during the next years. He is currently lunging and has been backed. He has shown very successfully locally in hand, being placed first and champion at Ashgar’s winter show as a three-year-old and first place at the Welsh Society’s Summer Show in Johannesburg in 2010. He will make his first national show debut at the Horse of the Year show in 2012.

Article written by: Zena Penfold (Millsleigh Irish Horse and Pony Stud) Contact details: info@dipereng.co.zazpenfold@iafrica.com , cell: +27 836161094

 

USA: Mountain Lad Line

THE MOUNTAIN LAD LINE
A REPORT FROM DONNA DUCKWORTH IN THE USA

In 1980 it was thought that the Mountain Lad stallion line, one of three major Connemara stallion lines, was in grave danger. The situation was thought to be so dire that some extraordinary measures to save the line were taken. In the US, May Medley in Georgia started to breed her son of Mon Tully Man (Tully Lad X Tooreen Pet [by Tooreen Laddie]), who she had been keeping with a gelding in her backyard for many years. Montully Son, the stallion, is a great grandson of Mountain Lad through his grandsire Tully Lad. May had been told by Caroline Nesbitt that he was the last of the Mountain Lad line. Montully Son was out of a daughter of Marconi, Marconi’s Stormy Rebel. May also bought a daughter of Jiminy Cricket, another son of Tully Lad, so she could line breed from the Mountain Lad line. This pony, Aillte Mhuire, had had one daughter by Custusha’s Cashel Rock, Courtney’s April Frolic, before coming to Georgia where she produced many offspring. Courtney’s April Frolic is the dam of the winner of the ACPS Clifden Trophy for the Purebred Connemara in the USEF Pony Hunter divisions who receives the most points in USEF sponsored competition, Balmullo’s Azalea who is shown as Ooh La La.

Aillte Muhire’s progeny by Montully Son were also highly successful. Tullymor’s April Fool was a beautiful and productive mare whose son, Balmullo’s Beacon (Tullymor’s April Fool X Aladdin), owned by Vanessa Morgan, was a successful eventer and prolific sire for Foothill’s Farm in Tennessee. May also bred Montully Son to many other what she called “old-fashioned” Connemaras, those that she could find with no Little Heaven blood in their pedigree. This program was in many ways extraordinarily successful with Tullymor ponies, including many stallions, now in every corner of the US, where they successfully compete in driving, eventing and dressage.

While all this was happening in the US, in England, a special consortium was formed to save the aged stallion Thunderbolt from the knackers. Thunderbolt was by Thunder, another son of Tully Lad and out of Irene Grey, who was a granddaughter of the Irish Draft, May Boy. This program has also produced ponies that are found far and wide. In Europe Chiltern Thunderburst (Thunderbolt x Chiltern Cameo) and his son, Rory Rasputin (Chiltern Thunderburst x Van de Aronbosch Donna) are very popular sires in Denmark and The Netherlands, respectively. In England there is Spinway Comet (Thunderbolt x Spinway Cailin) who is a many-times winner in hand. In the US, there is a son and a grandson of Thunderbolt in California. The son, Lasrachai, is out of Lovely Marcella (by Rebel Wind). The grandson is Sydserff Avalon (Cocum Camelot [by Thunderbolt] X Ballydonagh Misterina [by Kirtling Brigadoon]). There is another grandson in Indiana, Landsdown Willow’s Harbor Boy (*Chiltern Colm X Sillbridge Miranda [by Thunderbolt]). *Callowfeenish Thunder, Sam Davis’ famous gelding, Tor, was a Thunderbolt son imported to the US as a stallion and sired at least one pony in Virginia, Aluinn Torrence, before being gelded.

us_mountain_lad_lineIn 1983 in Ireland Tully Grey (Tully Lad X Cait Nu Dhuibhir) was bred to Abbeyleix Bluebird (Coosheen Finn X Blue Moon), starting a third major branch of the Mountain Lad line, the “Hazy” pony line, perhaps the most prolific of all the branches. The result of this cross, Dale Haze, has sons and grandsons throughout Europe and the US through the stallions Hazy Dawn, Moy Hazy Cove and later, Hazy Match. Fernville Matchmaker (Hazy Match X Ross Castle Moya [by Abbeyleix Owen]) was imported to the US two years ago and has gotten his first mare in foal.

Now the Mountain Lad genes are spread throughout the world and it is hard to find a Connemara pedigree without Mountain Lad in it somewhere. The importation of Frederiksminde Hazy Cavalier (Hazy Dawn X Frederiksminde Christina) to Australia assures that the line will continue to increase! Sue Clark of the Glenormiston stud in Australia says that Cavalier has filled out a lot since he arrived. Sue also said, “Unfortunately his dam Frederiksminde Christina died recently at the age of 25 so he is the last of this (Christina) line. His sire Hazy Dawn is also in his 20’s but will hopefully continue to breed for a few more years.

Connemara News and Thoughts in Ireland’s Equestrian

CONNEMARA NEWS – AND – THOUGHTS UPON WHICH TO PONDER

Those of you who are lucky enough to be able to purchase the magazine “Ireland’s Equestrian” will probably have already read the article on “Kippure Stud – The Business of Connemaras”

Many of you will probably have visited “Kippure”, spoken to Michael Igeo and realised the depth of Connemara knowledge that he possesses and will readily appreciate the message that is contained in his article on page 69 – 71 of the March April issue of Ireland’s Equestrian.

Michael outlines the primary use of the Connemara Pony in the early days and the market forces that saw change in the use of the Pony – consumerism, specialisation, dependence and market propaganda prevailed with the system of brood mares no longer expected to be worked and tested thoroughly. Faults and weaknesses that other wise would be found at the source of the working were now getting through. He speaks of a broad universal knowledge that is now found in place of what was a narrow detailed personal knowledge of individual ponies and the breed in general that lasted from childhood until death. Succession meant inherited knowledge and the passing on of this knowledge.

Understanding the relationship of “form and function” holds the key to success. Get it wrong and the result is devastating he states. Selective breeding is an art, which is second nature to someone who is gifted with the knowledge. Understanding the bloodlines and back breeding requires patience and analysis. He refers to the knowledge of Susan De Vesci, Jimmy Jones and Graham Tulloch that represented a Life’s work, a philosophy committed to memory.

In the heading – “Links in the chain” -Michael refers to the activities of the later part of the 20th century and the problems of inbreeding, over height, lack of bone etc. Michael states that as we delve further into the Connemara Pony, the more impressive is the impact that some of our predecessors have made. These vital links must be maintained, as without them, the demise is hastened.

“Need for reform –Blueprints for competition” Michael believes that our endeavours are to build on the efforts of our past stalwart breeders and be authentic in this restoration. The “In-Hand” world is only a small glimpse into the story of the Connemara Pony. Ponies must be bred that will meet customers expectations. There is a need to instil confidence in the industry of breeding Connemara ponies that in fact possess those unique qualities of the Connemara pony that can and do perform As well as and in fact better than most.

An interesting and thought provoking article for those who have concerns for the future of the Connemara pony.

Ireland: Fabulous Derby Win For Ballyowen Maybelle Molly

FABULOUS DERBY WIN FOR BALLYOWEN MAYBELLE MOLLY

The seven year old Irish Connemara mare, Ballyowen Maybelle Molly, competed at the Complet International de Fontainebleau Show Jump Championships in France on 18th and 19th April 2009.

Molly competed in the final seven year old International Jumping Competition over a huge, highly technical course. Molly had the only treble clear round out of the entire field. Huge congratulations must go to her rider, Kelly Allen and owner Anthony McCormac. The results of the mare’s two days competition are:

CSIP Table A against the clock, International Jumping Competition for 7 year old, FEI Art. 238.2.1 1m15 Saturday 18th April:

  1. BALLYOWEN MAYBELLE MOLLY (ISH); Kelly Allen (IRL) Faults 0.00 – time 34.99
  2. LITTLE DARK WARRIOR (IRRP) ; Niamh Dooley (IRL) Faults 0.00 – time 57.67
  3. OMBRE DES FALAISES; Robin Muhr (FRA) Faults 0.00 – time 65.15
  4. IRISH NIGHT WATCHMAN; Brian Crowley (IRL) Faults 0.00 – time 72.83

CSIP Pony final seven year old, International Jumping Competition Table A with jump off. FEI Art.238.1.2 1m20 Sunday 19th April 2009:

  1. BALLYOWEN MAYBELLE MOLLY (ISH); Kelly Allen (IRL) Faults 0.00/0.00 – time 34.28
  2. IRISH NIGHT WATCHMAN; Brian Crowley (IRL) Faults 0.00/4.00 – time 34.93
  3. GARRYDNUG GOLD DIGGER (ISH); Madeline Roberts Allen (IRL) Faults 0.00/4.00 – time34.99

A terrific result for owner, rider and the Connemara pony in general.bally

BALLYOWEN MAYBELLE MOLLY

It was a joyous occasion for all her connections – owner Anthony Mac Cormack, rider Kelly Allen and breeders John and Jenny Richardson when Ballyowen Maybelle Molly won the seven year old derby at Fontainbleau on April 19th 2009. She had become, officially, the best seven year old in Europe. Her magnificent achievement was not a surprise to me because I know the family and have followed its fortunes for many years

It all started when Sean Mac Lochlainn bought a three year old filly by Tiger Gill out of a Tully Lad mare called Kiss me Kate. She was a dun roan and stood at 13 hh. Sean entered her in Class14 at the Clifden Show in August 1960. This was a class for non registered Connemara mares not less than three years old. His sole purpose in doing so was that such ponies could be inspected for registration at the show which she was. She was called Lus na bPog (2103).

Her career as a show pony was short and inglorious but she became the most wonderful broodmare and left five fabulous daughters.

The oldest, Poigin (Carna Dun) went to England/Scotland. Ballyowen Belle (Carna Dun) stayed in Ireland with the Richardsons. Ciotog (Doon Paddy) went to Montana/Canada. Bla an Duinin (Doon Paddy) went to Sweden and Doireann (Macdara) went to France.

There is obviously a big Little Heaven influence but I also think there is a big female thing going on in this family The females are so dominant that we mustn’t give too much of the credit to the ‘topline’. Consider also that Doireann, who was by Macdara (no outside blood), produced Balladin de Fremur (Macky) in France, a registered stallion who jumped at national level there, thus garnering a big reputation for himself and the breed.

The extended Lus na bPog family is littered with great performers but three of them were genuine superstars operating at the very highest level. They were Poigin, Golden Avenns Ruby (daughter of Ciotog) and Molly (daughter of Belle). All three were females ridden by extremely talented girls – Fiona Clarke, Melanie Jacobi and Kelly Allen respectively.

Poigin and Goldie were allrounders, excelling in a variety of disciplines. They looked alike being typical of the Carna Dun line i.e. tall, scopey strong heads. Molly is different in that she is an absolute jumping machine and looks as well as she jumps. Doris Jacobi and I have written previously about Poigin and Goldie (English Chronicle 2001) and all I can say about Molly, being a seven year old, is that her career, stunning as it is to date, is only beginning.

The prominence of the females in the family is intriguing The talent and athleticism is unquestionable and if Sean were alive today we can assume that he would regard the £77-10-0 that he paid for Lus and her companion Londubh fifty years ago, as pretty good value!

Tom MacLochlainn

Blue Eyed Cream Ponies

BASE, SINGLE & DOUBLE DILUTE COAT COLOURS EXPLAINED

by Sorrel Lambton (BHSAI . ICES)

Fundamental Genetics

Horse coat colour genetics is an interesting, complicated, and sometimes confusing subject! As more research is carried out we gain a greater understanding as to how, why, and where some of the more unusual coat colours stem from. This is a simplified explanation, which I hope will dispel some of the myths surrounding foals born with unusual coat colours. A horse possesses 32 pairs of chromosomes in each cell, which carries unique information from generation to generation. Both the stallion and the mare’s genetic contribution is equal. When conception takes place genes are not lost or diluted in any way but combined. Horses can be described as either:

HETEROZYGOUS: means the horse possess two forms of a particular gene encoding some inheritable characteristic, which may therefore produce offspring differing from their parents and each other in that characteristic. These horses possess one masking gene (the dominant gene) and one hidden gene (the recessive gene). Because the recessive gene is hidden the horse will appear exactly like a homozygous horse.

HOMOZYGOUS: means the horse has two identical genes at the same place on two corresponding chromosomes. When a homozygous dominant horse is bred to a recessive horse (which by definition is always homozygous), all progeny will look like the homozygous dominant parent, but unlike the parent they will be a carrier (heterozygous) for the recessive gene.

The phenotype (outward coat colour appearance) can be the same with two horses but their genotype (genetic makeup and the information carried by the genes in the cells) can differ. Because the recessives are unknown it can be difficult to determine the outcome of coat colour. Unless we know the birth colour of the sire and dam and understand the reasons why a coat colour occurs the resulting foal colour will be difficult to determine. The only 100% reliable decision is to have a blood test or hair sample test to confirm the genetic makeup of each parent to eliminate guess work and so the ‘chance factor’.

Base colours (C+C+)

There are only two basic coat colours, black (fading and non fading), and red (chestnut). Black, bay, brown and chestnut are called base colours. A base coat colour can only be diluted to a single dilute (C+Ccr) to produce a buckskin (called dun in Ireland) or palomino if a diluting gene is introduced in the absence of other modifying genes. A horse of a base colour does not carry the diluting gene so can never produce a BEC. For example if you bred a BEC mare to a bay or brown stallion the foal will be buckskin. If the same BEC mare is bred to a chestnut the foal will be 100% palomino every time. In general lighter colours are dominant over darker colours. Red (chestnut) is the most recessive colour followed by black. Light bay is dominant over bay, which is dominant over brown, which in turn is dominant over black. All other coat colours are determined by the addition, alteration or absence of these base colours.

Sometimes a horse looks black but in reality is the darkest form of buckskin and so is classified as a single dilute. Unless this horse tests negative for the BEC gene you may have a surprise when breeding!

DILUTE COLOURS

Buckskin and Palomino are called single dilutes (C+Ccr) and each carry the diluting gene that can produce a BEC.

Dun (DD) and Buckskin (C+Ccr) compared

*The term dun used in Ireland is called a buckskin in other countries.

*There is a difference between the genetic makeup of a dun and a buckskin.
*A buckskin is denoted by the symbol C+Ccr and carries a single diluting gene where as a dun is denoted by the symbol DD and can never produce a BEC..
*Dun can be a breed as it can breed pure and will never produce a BEC.
*The dunning gene is dominant and affects all base colours without tending to lighten the legs or front of the face and leaves a darker mask.
*Duns have what are known as primitive markings. Failure to have a well defined dorsal stripe (a stripe along the spine extending into, and matching, the main and tail), and at least one other matching primitive marking like zebra marks on upper legs, or stripes or mottling on other parts of the body means it is not a D-genotype and more likely to be a buckskin (C+Ccr) and a carrier of the single dilute mechanism whichcan produce a BEC.
*Confusingly, some buckskins have a faint dorsal stripe – these are known as linebacked buckskins.
*Buckskins however never have primitive markings.
*Buckskin bred to buckskin or palomino, and some greys can produce a BEC.
*Buckskin is best bred to a base colour and will produce a buckskin, palomino, chestnut, bay, brown or black, depending on which base colour it is bred to – but will never breed a BEC.
*Buckskins bred from a palomino parent will often have some white hair at the base of the tail and sometimes in the main – known as frosting.
*Buckskin can never become a breed as it cannot breed true. It is only a coat colour.

Palomino genotype: C+Ccr

*A palomino like a buckskin is a single dilute. It occurs when a chestnut coat colour is diluted by the action of one diluting gene.
*A palomino should never be bred to another single dilute like buckskin or palomino.
*Palomino bred to grey can produce a BEC unless the grey tests negative for the diluting gene.

*Palomino bred to chestnut produces 50% palomino and 50% chestnut.
*Palomino bred to a base colour will produce similar coat colours like the buckskin but never a BEC. *Palomino can never become a breed for the same reasons as buckskin. It is only a coat colour.

Double dilutes, pseudo-albino CcrCcr

Blue eyed cream (BEC), cream, psuedo-albino, shiny eyes and cremello are some of the terms used in Ireland, and in other countries, to describe ponies and horses born with pinkish skin (sometimes called pumpkin), off white to cream coloured coats, and pale blue eyes that can appear pink in some light. Other less known terms include perlino, smoky cream or smoky perlino which all look similar except for subtle differences in colour due to the base colour the cream gene is acting on. The genetic mechanism (CcrCcr) which produces these coat colours is called a double dilute and can only occur when both the sire and the dam carry the Ccr gene. Sometimes the BEC has been called an albino which is completely incorrect as albinism does not exist in equines. A true albino has no skin/melanin pigment and can have hearing and sight problems. The term suedo-albino is a form of partial albinism which means they do posses pigment but not as much as other coat colours. A BEC’s eyes, just like a human with blue eyes, will be more light sensitive but no records that I have read so far report blindness. A worldwide survey was carried out in 2000 by the BEC Study Committee (email: mewatson@earthlink.net) via questionnaires to fifty-five connemara owners covering important questions like: light sensitivity, hearing, sight, melanoma, medical problems, rashes, hardiness etc. The results were nearly 100% positive with no one reporting any problems other than some of the BEC’s experiencing nose rash and slight light sensitivity with their pale eyes. Nose rash can occur from sunburn or sensitivity to irritants where any horse with a pink mussel is grazing.

Grey (GG)

Grey horses can be a very grey subject as the colour describes!
*Grey is not a true colour but rather an admixture of light and dark hairs superimposed over the horses entire body, which gradually lighten with age.
*The hair follicle of a grey horse is supposed to be, not only defective, but also not as deep-rooted in the dermis as other horses hair follicles, which is the reason why some people believe, they ‘grey out’, usually by the time they reach 6-8 years of age.
*Greys are more susceptible to melanomas, than other coat colours including BEC’s, of which 95% of cases are benign. However grey horses do suffer from gradual depigmentation usually around the eyes and mussel.
*Grey masks all other coat colours and causes many surprises in the horse-breeding world if the parents are not tested for the diluting gene.
*The birth colour of a foal whom is likely to turn grey should always be recorded as this is an indication of a foals true colour and will help determine how the horse will breed in the future. If for example a foal is born black, chestnut, bay or brown it should not carry the diluting gene and therefore will not breed a BEC foal. If on the other hand a foal is born buckskin it will carry the diluting gene and can breed a BEC foal if bred to another horse caring the dilute gene.
*If a grey shows speckled of fleabitten marks, (red speckles=chestnut; black=black; yellow=dun, buckskin, or palomino) in his coat, this is an indication of the true coat colour/birth coat colour and a guideline for breeders in choosing a correct mate.
*Every grey horse should be tested for the diluting gene CcrCcr to avoid the chance breeding of a BEC.

Conclusion

For the duration of the BEC ban, which spanned almost thirty years, the percentage of buckskins in the connemara breed has approximately halved, and the number of connemaras with grey coat colours has increased from approximately 50% to 70%. Black, chestnut, roan and palomino coat colours have reduced from approximately 10% to 3% since 1960.
Quote from Deidre Feely’s report of 2003:

‘The increase in the proportion of grey ponies in the population over the past few decades has been coupled by a decline in the percentage of dun, brown, black, roan and chestnut ponies. Thus, in recent years, the diversity amoung coat colour in registered Connemara Ponies has diminished’.
I have tried to be as accurate as possible in compiling and writing a simplified insight into how different connemara coat colours are achieved which I hope will give owners and breeders a greater understanding of how, or how not to, breed certain coat colours. There is a place for the BEC mare in breeding the popular buckskin (known as dun in Ireland), if the stallion is chosen wisely!

For anyone interested in exploring the subject further I suggest: *Horse colour explained by Jeanette Gower
*Wendy Bockman’s website www.doubledilute.com
*Equine Colour genetics by D.P. Sponenberg

*Coat colour genetics by Dr. Ann Bowling

*Coat Colour trends in the Connemara Pony Population in Ireland by Deirdre Feely B. Agr.Sc., Patrick Brophy MVB MRCVS., and Katherine Quinn M. Agr. Sc.
*Characterisation of the Connemara Pony Population in Ireland by Deirdre Feely B. Agr. Sc., Patrick Brophy MVB MRCVS., and Katherine Quinn M. Agr. Sc.

Test to detect the Blue Eyed Gene

It is now possible to have your connemara pony tested in Ireland for the BEC gene through Weatherbys Ireland DNA Laboratory at the Irish Equine Centre, Naas, Co. Kildare. The cost for this service is €30.00 per animal which will be significantly reduced pending negotiations with the Connemara Pony Breeders Society based on future testing of the entire foal crop. The test usually takes between two to three weeks but they will facilitate urgent requests if necessary. Simply ring Laura on 045-875521 or email her on: dnalab@weatherbys.ie with the breeding of your connemara pony.

Copyright Sorrel Lambton 2009

Characterisation of the Connemara Pony Population in Ireland

Report presented to
The Department of Agriculture, Food and Rural Development,
Agriculture House, Kildare St., Dublin 2.

Report funded by
The Department of Agriculture, Food and Rural Development,
Agriculture House, Kildare St., Dublin 2.

Deirdre Feely B.Agr.Sc.1,
Patrick Brophy MVB MRCVS1,
Katherine Quinn M.Agr.Sc1.

1Department of Animal Science and Production,
Faculty of Agriculture,
University College Dublin,
Belfield, Dublin 4.

Introduction

The Connemara Pony is numerically a small breed, with approximately 2,000 breeding females and 250 breeding males in Ireland. Traditionally, the Connemara was a working pony and enjoyed a prominent role in agricultural life in the West of Ireland. However, in the middle of the last century farming practices changed, and as machinery was introduced, the role of the working pony became redundant. The Connemara Pony has maintained its popularity by establishing a position in the showing and riding industry.

The Connemara Pony Breeders’ Society was founded in 1923. The main objectives of the Breed Society are the encouragement, development and maintenance of the Connemara Pony as a pure breed. Since its formation the Society has also been responsible for the publication of the Connemara Pony Stud Book. The breed is now recognised throughout the world as a top class performance pony and 17 different countries have established their own Breeders’ Societies.

As yet, there is little concern regarding the number of pure bred foals produced annually. However, the Connemara Pony Stud Book has been closed since 1964 and the practice of overusing popular sires is prevalent throughout the history of the breed. This could potentially led to a narrowing of the gene pool, high levels of inbreeding and a loss of genetic diversity within the population.

The main objectives of this project were to demographically and genetically characterise the Connemara Pony population, with specific emphasis placed on how past breeding practised have affected the present genetic composition of the breed. Height trends were also analysed in attempt to establish evidence of genetic erosion.

Methodology

A total of 20,032 records were used in the characterisation of the Connemara Pony population. These records were obtained from the Breed Societies’ database and from Dan-Axel Danielsson of the Swedish Connemara Pony Society.

The study focused on two reference populations. The first reference population consisted of 2,316 registered ponies born between 1993 and 1996 inclusive. This reference population represents the current breeding population of Connemara Ponies. The second reference population contained 2,844 foals born between 1998 and 2001 inclusive. This reference population contained records of both registered and non-registered ponies and represents the future breeding stock of the Connemara Pony. It should be noted that approximately one third of the non-registered ponies in this reference population will not be subsequently registered as Connemara Ponies.

The reference populations were characterised both demographically and genetically. The demographic characterisation is a description of a population in numerical terms and enables us to see how the size and structure of the population has altered over time. The main parameters estimated as part of the demographic characterisation included the number of registered ponies born year, the sex ratio, the average generation interval and family size. The genetic characterisation of a population is conducted to determine the level of genetic diversity within the population. As part of the genetic characterisation the average inbreeding coefficient and average relationship coefficient for animals in the reference populations were calculated. The contributions made by the ancestors of the reference populations, the number of founders, the effective number of founders and the effective number of ancestors were also estimated in order to measure the level of genetic diversity within the population. The influence that the Thoroughbred, Arab, Irish Draught and Welsh Cob breeds had on the reference populations was calculated. Height trends in the Connemara Pony were also analysed.

Summary of results

  • Demographic characterisation of the population
  • Up until 1959 the number of registered ponies born annually was consistently below 100. There was a steep increase in the number ponies born per year between 1959 and 1970, peaking at 568 in 1970. The population size was reduced considerably between 1971 and 1980, but recovered again between 1980 and 1996. At present there is approximately 800 pure bred foals born annually, however only 60 to 70% of these are subsequently registered as Connemara Ponies.
  • In recent years the ratio of registered mares to stallions deteriorated considerably. In 1980 for every 8.53 registered mares born there was one registered stallion born. In 1994 this ratio increased to 30.5 registered mares per stallion.

Demographic analysis of the reference populations

  • In the 1993 to 1996 reference population the number of registered ponies born per year increased from 548 in 1993, to 590 in 1996. In the 1998 to 2001 reference population the number of foals born per year appeared to decrease from 866 in 1998, to 441 in 2001. However, it is assumed that this is due to a lack of records for foals born in more recent years rather than an actual drop in annual foal production.
  • 181 different sires produced the 2,316 animals in the 1993 to 1996 reference population. The number of progeny per sire ranged from 1 to 140. 214 different stallions produced the 2,844 animals in the 1998 to 2001 reference population. The number of foals per stallion ranged from 1 to 151.

Age profile of the sires and dams of the reference populations

  • The average age of sires as higher for the more recent reference population. For example, 7% of the sires of the 1993 to 1996 reference population were under 5 years of age, but in the 1998 to 2001 reference population only 2% of the sires were under 5 years of age.
  • A larger proportion of younger dams produced the more recent reference population. 15% of the dams of the 1993 to 1996 reference population were 20 years or older, while only 5% of the dams of the 1998 to 2001 reference population were of that age group.


Generation interval

The generation interval is defined as the average age of the parents when their offspring are born.

  • The average generation interval between parents and offspring in the 1993 to 1996 reference population was 10.51 years. The average generation interval for the 1998 to 2001 reference population increased slightly to 10.59 years.
  • For both reference populations the generation interval between sires and their offspring was approximately 2 years longer than the generation interval between dams and their offspring.
  • Between 1980 and 2000 the average generation interval increased by 16%, from 8.98 years to 10.44 years.
  • The generation interval calculated is similar to the generation intervals found in other horse populations. The longer generation interval between sires and their progeny may indicate that breeders have a preference for older and proven sires, or it may merely reflect that stallions tend to commence breeding later in life.

Family size

For the purpose of the analysis family size was defined as the number of ‘breeding’ offspring per sire and dam. Offspring were deemed ‘breeding’ if they had produced at least one registered offspring themselves. In an ideal situation, family sizes would be balanced, giving each breeding animal an equal chance of producing their own replacements in the next generation.

  • On average each sire produced 2.88 ‘breeding’ male and 9.94 ‘breeding’ female offspring. Paternal family sizes were found to be extremely unbalanced, with a large proportion of the breeding population being produced by a small pool of stallions. For example, 10% of sires produced 55% of the ‘breeding’ female offspring and 30% of the ‘breeding’ male offspring.
  • On average each dam produced 1.24 ‘breeding’ male and 1.77 ‘breeding’ female offspring. The maternal family sizes showed less variation compared to paternal family sizes, as dams are biologically limited to producing one foal per year.
  • Paternal family sizes were very unbalanced. This is likely to cause a loss in the genetic variation of the breed, an increase in the relationship among animals in future generations, and a rise in the level of inbreeding.
  • Genetic characterisation of the population


Pedigree completeness

Pedigree completeness is an important parameter as the accuracy of the genetic characterisation is largely dependent on the quality of the records used in the analysis. Pedigree completeness was measured by determining the proportion of ancestors known per generation. The complete generation equivalent was also used to measure pedigree completeness and is defined as the average number of complete generations recorded.

  • Both reference populations’ pedigree data was practically 100% complete for the first 3 generations, i.e., almost 100% of all parents, grandparents and great grandparents were known. After the 5th and 6th generations, the proportion of known ancestors steadily decreased.
  • There complete generation equivalents for the 1993 to 1996, and the 1998 to 2001 reference populations, were 6.15 and 6.59 respectively.
  • The level of pedigree completeness for the animals in the reference populations was deemed to be relatively high when compared to corresponding studies.


Inbreeding

The coefficient of inbreeding measures the probability that an animal receives identical genes by descent from its sire and dam.

  • The average inbreeding coefficient for animals in the 1993 to 1996 reference population was 4.49%. This had increased to 4.65% for the 1998 to 2001 reference population.
  • 5.31% of the animals in the 1993 to 1996 reference population had inbreeding coefficients under 2%. The proportion of ponies in the 1998 to 2001 reference population with inbreeding coefficients under 2% was 2.67%.
  • The increase in inbreeding from 1980 to 1990 was 0.93% and was very similar to the expected rate at which inbreeding would increase under random mating conditions.
  • The average inbreeding coefficient for animals born in 1980 was 3.19%, this has increased steadily, reaching 4.65% in 2000.
  • The level of inbreeding detected in the Connemara Pony population was high in relation to most comparable studies. The actual increase in inbreeding corresponded to the theoretical increase in inbreeding expected if mating were at random, indicating that breeders did not take sufficient steps to avoid the mating of related animals.

Average relationship coefficient

The average relationship coefficient measures the proportion of genes that animals have in common.

  • The average relationship coefficient among animals in the 1993 to 1996, and 1998 to 2001 reference populations were 10.26% and 10.66% respectively. The relationship among the sires of both of the reference populations was almost 1% higher than the relationships among the dams of the reference populations.
  • Considering the average relationship between two first cousins is 12.5%, the average relationship among animals in the reference populations was extremely high. The average relationship among the sires of the reference populations was higher than the relationship among the dams of the reference populations, implying that stallions selected for breeding are of similar ancestry or breeding lines.

Contributions made by ancestors 

Important ancestors were identified by calculating the marginal contributions made by ascendants to the reference populations. The marginal contribution is the contribution made by an ancestor that is not already explained by another animal.

  • Carna Bobby was the most important ancestor to both of the reference populations, with a marginal contribution of 13.81% to the 1993 to 1996 reference population and 13.93% to the 1998 to 2001 reference population. Dun Lorenzo and Carna Dun were the next most important ancestors, contributing approximately 10.5% and 8.5% respectively to the genes of the reference populations.
  • Overall the contributions made by the ancestors of the reference populations were found to be very unequal with 6 ancestors contributing 50% of the genes to both of the reference populations
  • The imbalance of the contributions made by ancestors implies that future generations are at risk of further losses in genetic variation.

Number of founders, effective number of founders A founder is defined as an ancestor with unknown parents or the unknown parent where only one parent is unknown. It is assumed that all founders are unrelated and all of the genes in the populations emanate from these founders. The effective number of founders is a theoretical number defined as the number of equally contributing founders that would be expected, given the level of genetic diversity that exists in the reference population. The more balanced the founder contributions are to the reference population, the more the effective number of founders will approach the actual number of founders.

  • There were 351 founders for the 1993 to 1996 reference population and 342 founders for the 1998 to 2001 reference population.
  • The effective number of founders for both of the reference populations was 35.8.

The discrepancy between the actual number of founders and the effective number of founders is expected to decrease the amount of genetic diversity in the present population relative to what would have transpired had all founders contributed equally.

The effective number of ancestors

The effective number of ancestors is a theoretical number that supplies us with the minimum number of ancestors needed to explain the complete genetic diversity of the reference population. Unlike the effective number of founders, the effective number of ancestors accounts for bottlenecks in the pedigree. The closer the effective number of ancestors is to the effective number of founders, the smaller the impact past bottlenecks have had on the genetic diversity of the population.

  • The effective number of founders for both of the reference populations was approximately 18.
  • This indicates that past bottlenecks have adversely affected the genetic diversity of the animals in the reference populations.

The breed composition of the reference populationsA small number of Thoroughbred, Arab and Irish Draught stallions sired registered Connemara Ponies in the 1940s and 1950s. There were also two stallions in the pedigree file that were known to have Welsh Cob genes. The proportion of genes that the animals in the reference populations possessed, originating from these stallions, was estimated to obtain the influence that the Thoroughbred, Arab and Irish Draught had on the reference populations..

  • The Thoroughbred was the most influential of the foreign breeds, accounting for approximately 6% of the genes of the reference populations. The Arab, Irish Draught and Welsh Cob accounted for approximately 3.7%, 1.2% and 0.9% of the genes of the reference populations respectively. Approximately 88% of the genes in the animals in the reference population are assumed to be Connemara Pony.
  • The majority of the animals in the reference populations possessed at least some Welsh Cob, Thoroughbred and Arab genes. Approximately 50% of the animals in the reference populations had Irish Draught in their ancestry.

Height trends in the Connemara PonyThe traditional role of the Connemara Pony was as a versatile working animal. The main emphasis was on producing ponies with strength, hardiness, good bone and intelligence. However, as farming became increasingly mechanised during the middle of the last century, the role that the Connemara Pony had secured as a working animal began to disappear. The breed has survived by gaining a reputation as a performance animal and establishing a place in the showing and riding industry. In order to adapt to present market demands the breed is moving away from a traditional type to a ‘modern’ type of animal that is taller and lighter boned. As the breed moves away from the traditional type valuable genes may be lost, along with the characteristics that distinguish the Connemara Pony from other equine breeds. This process is known as genetic erosion and is a problem confronting many present-day equine breeds. Height trends in the Connemara Pony were analysed in an attempt to identify the occurrence of genetic erosion.

  • The average height at time of inspection of registered Connemara Ponies born in 1970 was 135.33cm. This increased steadily over the years, reaching 142.88cm in 1997.
  • Between 1972 and 1997 the average height of registered mares and stallions at the time of inspection increased by 8cm and 5.5cm respectively. Between 1975 and 1995, ponies bred outside Co. Galway were significantly taller than ponies bred inside Co. Galway (141.29cm +/- 4.56 versus 139.23cm +/- 5.16; P=0.0001). However, between 1991 and 2000 there was no difference in height between the two groups (142.49cm +/- 3.98 versus 142.38cm +/- 4.18; P=0.3834).
  • The analysis of height trends confirmed that Connemara Ponies are growing taller. It is assumed that this is a consequence of both improved environmental conditions and selection for taller ponies.

Conclusion

Following the analysis, it appears that the Connemara Pony breed is being confronted with two problems. Firstly, the survival of the traditional type of breed is under threat, and secondly, the genetic diversity of the breed is diminishing.

Today, the riding industry is an important outlet for Connemara Ponies. However, there is concern that this industry is instigating a shift from the traditional type of pony, to a taller, ‘modern’ type. The traditional type of Connemara Pony is perfectly adapted to the environment in which it developed and is completely distinct from other equine breeds. It may be necessary to safeguard against market forces inciting the disappearance of the traditional type of pony, which is a valuable national resource, and once lost can never be recovered.

The results generated from the characterisation of the Connemara Pony population indicate that past breeding practices have caused a significant loss in the breeds’ genetic diversity. To ensure that the genetic variation in the breed does not recede to a detrimental level, breeding policies need to be altered.

In future, it is vital that sire family sizes become more balanced, giving all stallions a better opportunity to breed their own replacements in the next generation. This would help to control the level of inbreeding and genetic diversity within the population.

The stallions used for breeding are closely related to each other and tend to be of similar ancestry or breeding lines. From a genetic diversity perspective it may be advantageous to have a pool of breeding stallions that are less related to each other to bestow a variety of genes to the proceeding generations.

As relationships among animals in the present population is high the mating of related animals is inevitable. Breeders must be very vigilant in respect to the stallions that they use for breeding to ensure that a minimal amount of inbreeding is practised.

There are 16 different countries, outside Ireland, that have formed their own Breeders’ Societies and maintain their own stud books. A study is presently being undertaken to characterise the Connemara Pony populations in a number of these countries. It is hoped that these animals may be a source of genetic variability that could be used to widen the gene pool of the Irish Connemara Pony population.

Adequate genetic diversity is vital for the long term health and viability of any population. Thus, it is vital that breeding practices are altered in order to secure the future prosperity of the Connemara Pony breed.